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Hyperlane is a pioneering interoperability layer designed to connect the modular blockchain ecosystem, enabling seamless communication across diverse blockchain environments such as Layer 1s, rollups, and app-chains. Built with a focus on permissionless deployment, Hyperlane empowers developers to bridge chains effortlessly while offering customizable security models tailored to specific application needs. This flexibility makes it a cornerstone for creating interchain applications that operate across multiple blockchain networks, fostering a more interconnected and efficient Web3 landscape. Hyperlane’s architecture is centered around a smart contract 'mailbox' deployed on each supported chain, functioning as an onchain API for dispatching and receiving messages. This mailbox system allows for secure and efficient cross-chain communication, with successful message transfers requiring two transactions: one to send from the origin chain and another to deliver on the destination chain. Developers can enhance security by overriding the default Interchain Security Module (ISM) with application-specific configurations, ensuring that interchain messages are verified as authentic.